In 1980, Dizhou Tong, also called Ti Chou Tung, and his colleagues in China isolated nuclei from the embryonic cells of Cyprinus carpio Linnaeus, or Asian carp, and injected them into European carp eggs, for which the nuclei had been removed.

About ten percent of the clones developed into juvenile fish that had the traits of Asian carp and of European carp. This results was Tong’s last, and it was published in 1980, a year after his death.